Sorry, I am with Zenphoto since 2007 and I really don't remember that type of link ever being used or worked. If it did work it might been an unofficial way. My colleague may correct me.
Use the image template functions available instead of i.php directly:
http://www.zenphoto.org/documentation/functions/_template-functions.php.html
get versions get the url, print versions print the html tag.
Update: I stand corrected. You are indeed right, those links did once work and got lost with the mod_rewrite restructuring in 1.4.5. It is not done via hatchess directly anymore to better support servers without htaccess like nginx (which our own server uses).