thanks acrylian for the prompt reply.
may i request if someone can guide as i searched through 100's of pages in few hours but couldnt found matching htaccess for my site
www.wildart.in
wanted to convert
http://www.wildart.in/Woodpeckers%20and%20allies/
TO
http://www.wildart.in/Woodpeckers_and_allies/
Then
http://www.wildart.in/Woodpeckers%20and%20allies/Yellow-crowned%20Woodpecker/
TO
http://www.wildart.in/Woodpeckers_and_allies/Yellow-crowned_Woodpecker/
would really appreciate if someone can show me RewriteRule
Doing this is really important as i do upload my folder structure and images through gallery based on international bird scientific information and nomenclature is set based on that so i can't really change spaces to _ in real folder.
sorry for being too basic. am just a photographer setting up my Bird gallery with some information data in it.